Edited by Ykä Helariutta, University of Cambridge, Cambridge, United Kingdom, and accepted by Editorial Board Member Philip N. Benfey February 18, 2021 (received for review November 29, 2020)Plant miRNAs can move from cell to cell through cytoplasmic channels called plasmodesmata, transmitting positional information that is decoded to make developmental decisions; one well-studied example of this is xylem patterning in the root of the model plant Arabidopsis thaliana, which depends on the formation of opposing gradients of two miRNAs and their target transcripts. Here, we show that two...…Edited by Ykä Helariutta, University of Cambridge, Cambridge, United Kingdom, and accepted by Editorial Board Member Philip N. Benfey February 18, 2021 (received for review November 29, 2020)Plant miRNAs can move from cell to cell through cytoplasmic channels called plasmodesmata, transmitting positional information that is decoded to make developmental decisions; one well-studied example of this is xylem patterning in the root of the model plant Arabidopsis thaliana, which depends on the formation of opposing gradients of two miRNAs and their target transcripts. Here, we show that two...WW…